k Museum of Jurassic Marine Life, BH20 5PE Kimmeridge, United Kingdom.jSchool of Geosciences, University of the Witwatersrand, 2000 Johannesburg, South Africa.iDepartment of Biological Sciences, George Washington University, Washington, DC 20052.hEvolutionary Studies Institute, University of the Witwatersrand, 2000 Johannesburg, South Africa.gDepartment of Earth and Environmental Sciences, University of Iowa, Iowa City, IA 52242.fCentre de Recherche sur la Paléobiodiversité et les Paléoenvironnements, Muséum National d'Histoire Naturelle, 75005 Paris, France.eConsejo Nacional de Investigaciones Científicas y Técnicas, División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, National University of La Plata, B1900 La Plata, Buenos Aires, Argentina.dDepartment of Biomedical Sciences, Heritage College of Osteopathic Medicine, Ohio University, Athens, OH 45701.